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Sequencing Primers


Though Addgene mainly uses next-generation sequencing (NGS) for plasmid verification, Addgene has used a number of primers for Sanger sequence verification over the years. Below is a list of popular universal primers available from most sequencing services, along with a more complete list of common sequencing primers. These lists are provided for your reference, Addgene does not distribute primers.

To identify primers that may be useful in your sequencing reaction, find your plasmid’s page and see what primers are listed under "5' sequencing primer" and "3' sequencing primer" or explore the sequence map in SnapGene or other software to see additional annotated primers. You can also use Addgene’s protocol to design your own primers. Still have questions about choosing the best primer for your plasmid? Email us at [email protected].

For additional information on molecular biology, plasmids, and recombinant DNA, please consult Addgene's Molecular Biology Reference page.

Universal Sequencing Primers

Name Sequence (5' to 3') Description Direction
BGH-R TAGAAGGCACAGTCGAGG Bovine growth hormone terminator Reverse
CMV Forward CGCAAATGGGCGGTAGGCGTG Human CMV immediate early promoter Forward
EGFP-C CATGGTCCTGCTGGAGTTCGTG 3' end of EGFP Forward
EGFP-N CGTCGCCGTCCAGCTCGACCAG 5' end of EGFP Reverse
hU6-F GGGAAACGCCTGGTATCTTT Human U6 promoter Forward
LKO.1 5' (U6) GACTATCATATGCTTACCGT Human U6 promoter Forward
M13 Forward (-21) TGTAAAACGACGGCCAGT In lacZ gene Forward
M13 Reverse CAGGAAACAGCTATGAC In lacZ gene Reverse
M13/pUC Forward CCCAGTCACGACGTTGTAAAACG In lacZ gene Foward
M13/pUC Reverse AGCGGATAACAATTTCACACAGG In lacZ gene Reverse
MSCV Forward CCCTTGAACCTCCTCGTTCGACC Murine stem cell virus Forward
MSCV Reverse CAGCGGGGCTGCTAAAGCGCATGC Murine stem cell virus Reverse
pBABE 5' CTTTATCCAGCCCTCAC Psi packaging signal, 5' of MCS in pBABE vectors Forward
pBAD Forward ATGCCATAGCATTTTTATCC For vectors with E. coli araBAD promoter Forward
pBAD Reverse GATTTAATCTGTATCAGG For vectors with E. coli araBAD promoter Reverse
pBluescript KS TCGAGGTCGACGGTATC For pBluescript vector Forward
pBluescript SK TCTAGAACTAGTGGATC For pBluescript vector Reverse
pGEX 3' CCGGGAGCTGCATGTGTCAGAGG 3' end of MCS in pGEX vectors Reverse
pGEX 5' GGGCTGGCAAGCCACGTTTGGTG 3' end of glutathione-S-transferase Forward
SP6 ATTTAGGTGACACTATAG SP6 promoter Forward
T3 GCAATTAACCCTCACTAAAGG T3 promoter Forward
T7 TAATACGACTCACTATAGGG T7 promoter Forward
T7 Terminal GCTAGTTATTGCTCAGCGG T7 terminator Reverse

Other Common Sequencing Primers

Name Sequence (5' to 3') Description Direction
3'AOX1 GCAAATGGCATTCTGACATCC For Pichia vectors with AOX1 terminator Reverse
5'AOX1 GACTGGTTCCAATTGACAAGC For Pichia vectors with AOX1 promoter Forward
35S promoter CTATCCTTCGCAAGACCCTTC CaMV 35S promoter Forward
AC5 ACACAAAGCCGCTCCATCAG Drosophila Actin 5C promoer Forward
Alpha-factor TACTATTGCCAGCATTGCTGC Alpha factor signal sequence Forward
Amp-R ATAATACCGCGCCACATAGC 5' end of ampicillin resistance gene Reverse
AUG1 Forward CAATTTACATCTTTATTTATTAACG For Pichia vectors with AUG1 promoter Forward
AUG1 Reverse GAAGAGAAAAACATTAGTTGGC For Pichia vectors with AUG1 promoter Reverse
BGH-R TAGAAGGCACAGTCGAGG Bovine growth hormone terminator Reverse
Bglob-intron-F CTGGTCATCATCCTGCCTTT Rabbit beta-globin intron Forward
Bglob-intron-R TTTGCCCCCTCCATATAACA Rabbit beta-globin intron Reverse
Bglob-pA-R TTTTGGCAGAGGGAAAAAGA Rabbit beta-globin polyA region Reverse
CAT-R GCAACTGACTGAAATGCCTC 5' end of chloramphenicol resistance gene Reverse
CMV Forward CGCAAATGGGCGGTAGGCGTG Human CMV immediate early promoter Forward
CRE-R GCAAACGGACAGAAGCATTT 5' end of Cre recombinase Reverse
CYC1 GCGTGAATGTAAGCGTGAC CYC1 transcriptioin termination signal/td> Reverse
DsRed1-C AGCTGGACATCACCTCCCACAACG 3' end of DsRed1 Forward
DsRed1-N GTACTGGAACTGGGGGGACAG 5' end of DsRed1 Reverse
EBV Reverse GTGGTTTGTCCAAACTCATC SV40 polyA terminator Reverse
Ecdysone Forward CTCTGAATACTTTCAACAAGTTAC Drosophila heat shock promoter Forward
EF-1α Forward TCAAGCCTCAGACAGTGGTTC Human elongation factor-1α promoter Forward
EGFP-C CATGGTCCTGCTGGAGTTCGTG 3' end of EGFP Forward
EGFP-N CGTCGCCGTCCAGCTCGACCAG 5' end of EGFP Reverse
EXFP-R GTCTTGTAGTTGCCGTCGTC For distinguishing EGFP vs ECFP vs EYFP Reverse
F1ori-F GTGGACTCTTGTTCCAAACTGG F1 origin Forward
F1ori-R AGGGAAGAAAGCGAAAGGAG F1 origin Reverse
GAL1 AATATACCTCTATACTTTAACGTC S. cerevisiae GAL1 promoter Forward
Gal10pro-F GGTGGTAATGCCATGTAATATG S. cerevisiae GAL10 promoter Forward
Gal4 N-term GAGTAGTAACAAAGGTCAA 3' end of Gal4 DNA binding domain Forward
Gal4-AD AATACCACTACAATGGAT 3' end of Gal4 activation domain Forward
GFP-F GGTCCTTCTTGAGTTTGTAAC 3' end of GFP Forward
GFP-R CCATCTAATTCAACAAGAATTGGGACAAC 5' end of GFP Reverse
GPDpro-F CGGTAGGTATTGATTGTAATTCTG S. cerevisiae GPD promoter Forward
GW-3' GCATGATGACCACCGATATG 3' end of Gateway cassette Forward
GW-5' AATCTCGCCGGATCCTAACT 5' end of Gateway cassette Reverse
H1 TCGCTATGTGTTCTGGGAAA Human H1 promoter Forward
HA-F TACCCATACGACGTCCCAGA HA tag Forward
HA-R TCTGGGACGTCGTATGGGTA HA tag Reverse
HAT GAGGAGCACGCTCATGCCCAC Histidine affinity tag Forward
hGH-PA-R CCAGCTTGGTTCCCAATAGA Human growth hormone terminator Reverse
hU6-F GGGAAACGCCTGGTATCTTT Human U6 promoter Forward
hUBCpro-F TGAAGCTCCGGTTTTGAACT Human Ubiquitin C (UbC) promoter Forward
IRES-F TGGCTCTCCTCAAGCGTATT 3' end of IRES Forward
IRES-R CCTCACATTGCCAAAAGACG 5' end of IRES Reverse
L4440 AGCGAGTCAGTGAGCGAG 5' of MCS in L4440 vector Forward
LacI-R GGCATACTCTGCGACATCGT 5' end of LacI Reverse
LacZ-R GACAGTATCGGCCTCAGGAA 5' end of LacZ Reverse
LexA CGTCAGCAGAGCTTCACCATTG 3' end of LexA DNA binding domain Forward
LKO.1 5' (U6) GACTATCATATGCTTACCGT Human U6 promoter Forward
LNCX AGCTCGTTTAGTGAACCGTCAGATC Human CMV promoter Forward
Luc-F AGTCAAGTAACAACCGCGA 3' end of luciferase Forward
LucNrev CCTTATGCAGTTGCTCTCC 5' end of luciferase Reverse
M13 Forward (-21) TGTAAAACGACGGCCAGT In lacZ gene Forward
M13 Forward (-40) GTTTTCCCAGTCACGAC In lacZ gene Forward
M13 Reverse CAGGAAACAGCTATGAC In lacZ gene Reverse
M13/pUC Forward CCCAGTCACGACGTTGTAAAACG In lacZ gene Forward
M13/pUC Reverse AGCGGATAACAATTTCACACAGG In lacZ gene Reverse
MBP-F GATGAAGCCCTGAAAGACGCGCAG 3' end of maltose binding protein Forward
mCherry-F CCCCGTAATGCAGAAGAAGA 3' end of mCherry Forward
mCherry-R TTGGTCACCTTCAGCTTGG 5' end of mCherry Reverse
MT Forward CATCTCAGTGCAACTAAA Drosophila metallothionein promoter Forward
MMLV-F ATCAGTTCGCTTCTCGCTTC Moloney murine leukemia virus (MoMuLV) LTR Forward
mPGK-F CATTCTGCACGCTTCAAAAG Mouse PGK promoter Forward
MSCV CCCTTGAACCTCCTCGTTCGACC Murine stem cell virus Forward
MSCV-rev CAGCGGGGCTGCTAAAGCGCATGC Murine stem cell virus Reverse
MT1-F GCTGTCCTCTAAGCGTCACC Mouse metallothionein 1 promoter Forward
mU6-F CAGCACAAAAGGAAACTCACC Mouse U6 promoter Forward
Myc GCATCAATGCAGAAGCTGATCTCA Myc tag Forward
Neo-F CGTTGGCTACCCGTGATATT 3' end of neomycin resistance gene Forward
Neo-R GCCCAGTCATAGCCGAATAG 5' end of neomycin resistance gene Reverse
NOS-F GCGTTCAAAAGTCGCCTAAG Nopaline synthase promoter Forward
Nmt1-F GCAATGTGCAGCGAAACTAA S. pombe nmt1 promoter Forward
OpIE2 Forward CGCAACGATCTGGTAAACAC OpIE2 promoter Forward
pACYC-F TGAAGTCAGCCCCATACGAT p15A origin Forward
pAd-CMV GCTAGAGATCTGGTACCGTC For cloning sites after SalI in pAd-CMV vector Forward
pBABE 3' ACCCTAACTGACACACATTCC SV40 enhancer, 3' of MCS in pBABE vectors Reverse
pBABE 5' CTTTATCCAGCCCTCAC Psi packaging signal, 5' of MCS in pBABE vectors Forward
pBAD Forward ATGCCATAGCATTTTTATCC For vectors with E. coli araBAD promoter Forward
pBAD Reverse GATTTAATCTGTATCAGG For vectors with E. coli araBAD promoter Reverse
pBluescriptKS TCGAGGTCGACGGTATC For pBluescript vector Forward
pBluescriptSK TCTAGAACTAGTGGATC For pBluescript vector Reverse
pBMN 5' GCTTGGATACACGCCGC
MMLV sequence, for inserts in pBMN retroviral vector
MMLV sequence, for inserts in pBMN retroviral vector Forward
pBR322ori-F GGGAAACGCCTGGTATCTTT pBRS322 origin Forward
pBRforBam CTTGGAGCCACTATCGAC In pBR322 tet region, upstream of BamHI site Forward
pBRforEco AATAGGCGTATCACGAGGC In pBR322, upstream of EcoRI site Forward
pBRrevBam GGTGATGTCGGCGATATAGG In pBR322 tet region, downstream of BamHI site Reverse
pCAG-F GCAACGTGCTGGTTATTGTG Rabbit beta-globin intron, for pCAG plasmids Forward
pCasper-F GGGTTTTATTAACTTACAT 5' end of Drosophila mini-white gene Reverse
pCasper-hs GCAACTACTGAAATCTGCCAAG Drosophila Hsp70 promoter Forward
pcDL-F GTTGCCTTTACTTCTAGGCCT 5' of EcoRI site in pcDL vector Forward
pENTR-F CTACAAACTCTTCCTGTTAGTTAG 5' of attL1 in pENTR vector Forward
pENTR-R ATGGCTCATAACACCCCTTG 3' of attL2 in pENTR vector/td> Reverse
pGEX 3' CCGGGAGCTGCATGTGTCAGAGG 3' of MCS in pGEX vectors Reverse
pGEX 5' GGGCTGGCAAGCCACGTTTGGTG 3' end of glutathione-S-transferase Forward
pGP704-R AACAAGCCAGGGATGTAACG R6K gamma origin, 3' of MCS in pGP704 vector Reverse
pHybLex Reverse GAGTCACTTTAAAATTTGTATACAC ADH terminator Reverse
pLTet-F ACTGAGCACATCAGCAGGAC Lambda phage early leftward (pL) promoter Forward
pLXSN 5' (MSCV) CCCTTGAACCTCCTCGTTCGACC Murine stem cell virus, same as MSCV Forward
pMRB101-F AAGATGCAGGCAGCTGAGTT HCMV major immediate-early protein (IE) Forward
pMT2-F TTGCCTTTCTCTCCACAGGT 3' end of synthetic intron Forward
pMX-S1811 GACGGCATCGCAGCTTGGATACAC MMLV sequence, 5' of MCS in pMXs vector Forward
Polyhedrin forward AAATGATAACCATCTCGC Polyhedrin promoter Forward
Polyhedrin reverse GTCCAAGTTTCCCTG For baculovirus vector with polyhedrin promoter Reverse
pQE promoter CCCGAAAAGTGCCACCTG 5' of MCS in pQE vectors Forward
pREP Forward GCTCGATACAATAAACGCC Rous sarcoma virus (RSV) promoter Forward
pRS-marker CGGCATCAGAGCAGATTGTA To sequence yeast selectable marker in pRS vectors Forward
Pry1 CTTAGCATGTCCGTGGGGTTTGAAT PZ P-element Reverse
pTrcHis Forward GAGGTATATATTAATGTATCG 5' of MCS in pTrcHis vector Forward
pTrcHis Reverse GATTTAATCTGTATCAGG 3' of MCS in pTrcHis vector, same as pBAD-R Reverse
Puro-F GCAACCTCCCCTTCTACGAGC 3' end of puromycin resistance gene Forward
Puro-R GTGGGCTTGTACTCGGTCAT 5' end of puromycin resistance gene Reverse
pZIP TCCTTTCCAGCGAGGTTCTA Murine leukemia virus (MuLV) Reverse
RCAS-F ACATGGGTGGTGGTATAGCGCTTGCG 3' of Rous sarcoma virus (RSV) env gene Forward
Rluc-F CCAGGATTCTTTTCCAATGC 3' end of Renilla luciferase Forward
RVprimer3 CTAGCAAAATAGGCTGTCCC 5' of MCS in pGL3 vector Forward
SFFV-F ATTGATTGACTGCCCACCTC Spleen focus forming virus 5' LTR Forward
SP6 ATTTAGGTGACACTATAG SP6 promoter Forward
SV40pA-R GAAATTTGTGATGCTATTGC SV40 polyA Reverse
SV40pro-F TATTTATGCAGAGGCCGAGG SV40 promoter/origin Forward
SV40-spliceR CACAAAGATCCGGACCAAAG SV40 splice sequence Reverse
T3 GCAATTAACCCTCACTAAAGG T3 promoter Forward
T7 TAATACGACTCACTATAGGG T7 promoter Forward
T7 Terminal GCTAGTTATTGCTCAGCGG T7 terminator Reverse
Tac promoter GAGCGGATAACAATTTCACACAGG Tac promoter Forward
tdTomato-Fwd CTGTTCCTGTACGGCATGG 3' end of tdTomato Forward
tdTomato-Rev TCTTTGATGACGGCCATGT 5' end of tdTomato Reverse
Tet-R GGCGAGTTTACGGGTTGTTA 5' end of tetracycline resistance gene Reverse
TK-pA-R TTGTCTCCTTCCGTGTTTCA Thymidine kinase polyA Reverse
Tn7-end GGGGTGGAAATGGAGTTTTT Bacteria transposon Tn7 Reverse?
TRC-F CAAGGCTGTTAGAGAGATAATTGGA Human U6 promoter Forward
Ubx-F AACTCGTACTTTGAACAGGC Drosophila Ultrabithorax gene Forward
V5 Reverse ACCGAGGAGAGGGTTAGGGAT V5 epitope Reverse
WPRE-R CATAGCGTAAAAGGAGCAACA 5' end of WPRE Reverse
XBG-R GACTCCATTCGGGTGTTC Xenopus beta-globin 3'UTR Reverse
XEF1a TTTCGCCCTAACTTCGTGAT Xenopus EF1 alpha enhancer/promoter Forward
Xpress Forward TATGGCTAGCATGACTGGT Xpress epitope Forward

Content last reviewed: 22 October 2025