Skip to main content

A molecular switch for coordinating kinesin and dynein transport of mitochondrial cargo.

Gladkova C, Paez-Segala MG, Grant WP, Kittisopikul M, Myers SA, Wang Y, Vale RD
Science. 2026 Aug 6;393(6811):601-606. doi: 10.1126/science.aeh1475. Epub 2026 Aug 6. (Link opens in a new window) PubMed (Link opens in a new window) Article

Plasmids from Article

ID Plasmid Purpose
248233pHR Miro1 (1-590) m-sfGFP FLAG I3-01Encodes the cytosolic portion of human Miro-1 (RHOT1) C-terminally fused with monomerized sfGFP (V206R), FLAG and the self-assembling I3-01 domain.
248234pHR TRAK1 (1-702) mScarlet Encodes TRAK1 (1-702) C-terminally fused with mScarlet. Expression of this fragment facilitates TRAK1-mediated motility of Miro-1 decorated nanocages in cells.
248235pHR TRAK2 (1-700) mScarlet Encodes TRAK2 (1-700) C-terminally fused with mScarlet. Expression of this fragment facilitates TRAK2-mediated motility of Miro-1 decorated nanocages in cells.
248236pHR Miro1 (1-590) Halo FLAG I3-01Encodes the cytosolic portion of human Miro-1 (RHOT1) C-terminally fused with monomerized Halo, FLAG and the self-assembling I3-01 domain.
248237pHR TRAK1 E242R (1-702) mScarlet Encodes TRAK1 E242R (1-702) C-terminally fused with mScarlet. The E242R mutation disrupts kinesin-mediated transport.
248238pHR TRAK2 E242R (1-700) mScarlet Encodes TRAK2 E242R (1-700) C-terminally fused with mScarlet. The E242R mutation disrupts kinesin-mediated transport.
248239pHR TRAK1 L92A, F95A (1-702) mScarlet Encodes TRAK1 L92A, F95A (1-702) C-terminally fused with mScarlet. The L92A, F95A mutation disrupts the dynein regulatory helix (DRH).
248240pHR TRAK2 F92A, M95A (1-700) mScarlet Encodes TRAK2 F92A, M95A (1-700) C-terminally fused with mScarlet. The F92A, M95A mutation disrupts the dynein regulatory helix (DRH).
248241pHR TRAK2 A129V, A130V (1-700) mScarlet Encodes TRAK2 A129V, A130V (1-700) C-terminally fused with mScarlet. The CC1-box A129V, A130V mutations disrupts DLIC association.
248242pHR TRAK2 d373-380 (1-700) mScarlet Encodes TRAK2 d373-380 (1-700) C-terminally fused with mScarlet. The Spindly motif 373-380 deletion disrupts dynactin cofactor association.
248243pHR TRAK1 A129V, A130V (1-702) mScarlet Encodes TRAK1 A129V, A130V (1-702) C-terminally fused with mScarlet. The CC1-box A129V, A130V mutations disrupts DLIC association.
248244pHR TRAK1 d373-380 (1-702) mScarlet Encodes TRAK1 d373-380 (1-702) C-terminally fused with mScarlet. The Spindly motif 373-380 deletion disrupts dynactin cofactor association.
248245pHR TRAK2 S84A (1-700) mScarlet Encodes TRAK2 S84A (1-700) C-terminally fused with mScarlet. The S84A mutation prevents S84 phosphorylation.
248246pHR TRAK2 S84E (1-700) mScarlet Encodes TRAK2 S84E (1-700) C-terminally fused with mScarlet. The S84E mutation mimics S84 phosphorylation.
248247pHR TRAK1 (1-702) mScarlet, resi 81-106 from TRAK2Encodes TRAK1 (1-702) C-terminally fused with mScarlet. Residues 81-106 corresponding to the dynein regulatory helix are from TRAK2.
248248pHR TRAK2 (1-700) mScarlet, resi 81-106 from TRAK1Encodes TRAK2 (1-700) C-terminally fused with mScarlet. Residues 81-106 corresponding to the dynein regulatory helix are from TRAK1.
248249pHR TRAK1 L92A, F95A, d373-380 (1-702) mScarlet Encodes TRAK1 L92A, F95A, d373-380 (1-702) C-terminally fused with mScarlet.
248250pHR TRAK2 F92A, M95A, d373-380 (1-700) mScarlet Encodes TRAK2 F92A, M95A, d373-380 (1-700) C-terminally fused with mScarlet.
248251pHR TRAK1 T84E (1-702) mScarlet Encodes TRAK1 T84E (1-702) C-terminally fused with mScarlet. The T84E mutation mimics S84 phosphorylation.
248252pHR TRAK1 T84A (1-702) mScarlet Encodes TRAK1 T84A (1-702) C-terminally fused with mScarlet. The T84A mutation prevents T84 phosphorylation.
248253pHR TRAK1 (1-702) mScarlet FKBPEncodes TRAK1 (1-702) C-terminally fused with mScarlet and FKBP. Expression of this fragment facilitates TRAK1-mediated motility of FRB decorated nanocages in cells in the presence of rapamycin.
248254pHR TRAK2 (1-700) mScarlet FKBPEncodes TRAK2 (1-700) C-terminally fused with mScarlet and FKBP. Expression of this fragment facilitates TRAK2-mediated motility of FRB decorated nanocages in cells in the presence of rapamycin.
248255pHR TRAK1 L92A, F95A (1-702) mScarlet FKBPEncodes TRAK1 L92A, F95A (1-702) C-terminally fused with mScarlet and FKBP. The L92A, F95A mutation disrupts the dynein regulatory helix (DRH).
248256pHR TRAK1 A129V, A130V (1-702) mScarlet FKBPEncodes TRAK1 A129V, A130V (1-702) C-terminally fused with mScarlet and FKBP. The CC1-box A129V, A130V mutations disrupts DLIC association.
248257pHR TRAK1 d373-380 (1-702) mScarlet FKBPEncodes TRAK1 d373-380 (1-702) C-terminally fused with mScarlet and FKBP. The Spindly motif 373-380 deletion disrupts dynactin cofactor association.
248258pHR TRAK1 L92A, F95A, d373-380 (1-702) mScarlet FKBPEncodes TRAK1 L92A, F95A, d373-380 (1-702) C-terminally fused with mScarlet and FKBP.
248259pHR TRAK1 (1-702) mScarlet FKBP, resi 81-106 from TRAK2Encodes TRAK1 (1-702) C-terminally fused with mScarlet and FKBP. Residues 81-106 corresponding to the dynein regulatory helix are from TRAK2.
248260pHR TRAK2 F92A, M95A (1-700) mScarlet FKBPEncodes TRAK2 F92A, M95A (1-700) C-terminally fused with mScarlet and FKBP. The F92A, M95A mutation disrupts the dynein regulatory helix (DRH).
248261pHR TRAK2 A129V, A130V (1-700) mScarlet FKBPEncodes TRAK2 A129V, A130V (1-700) C-terminally fused with mScarlet and FKBP. The CC1-box A129V, A130V mutations disrupts DLIC association.
248262pHR TRAK2 d373-380 (1-700) mScarlet FKBPEncodes TRAK2 d373-380 (1-700) C-terminally fused with mScarlet and FKBP. The Spindly motif 373-380 deletion disrupts dynactin cofactor association.
248263pHR TRAK2 S84A (1-700) mScarlet FKBPEncodes TRAK2 S84A (1-700) C-terminally fused with mScarlet and FKBP. The S84A mutation prevents S84 phosphorylation.
248264pHR TRAK2 S84E (1-700) mScarlet FKBPEncodes TRAK2 S84E (1-700) C-terminally fused with mScarlet and FKBP. The S84E mutation mimics S84 phosphorylation.
248265pHR TRAK2 F92A, M95A, d373-380 (1-700) mScarlet FKBPEncodes TRAK2 F92A, M95A, d373-380 (1-700) C-terminally fused with mScarlet and FKBP.
248266pHR TRAK2 (1-700) mScarlet FKBP, resi 81-106 from TRAK1Encodes TRAK2 (1-700) C-terminally fused with mScarlet and FKBP. Residues 81-106 corresponding to the dynein regulatory helix are from TRAK1.
248267pHR TRAK1 fl mScarlet Encodes TRAK1 fl C-terminally fused with mScarlet.
248268pHR TRAK2 fl mScarlet Encodes TRAK2 fl C-terminally fused with mScarlet.
248269pHR TRAK1 L92A, F95A fl mScarlet Encodes TRAK1 L92A, F95A fl C-terminally fused with mScarlet. The L92A, F95A mutation disrupts the dynein regulatory helix (DRH).
248270pHR TRAK1 L92A, F95A, d373-380 fl mScarlet Encodes TRAK1 L92A, F95A, d373-380 fl C-terminally fused with mScarlet.
248271pHR TRAK2 F92A, M95A fl mScarlet Encodes TRAK2 F92A, M95A fl C-terminally fused with mScarlet. The F92A, M95A mutation disrupts the dynein regulatory helix (DRH).
248272pHR TRAK2 S84A fl mScarlet Encodes TRAK2 S84A fl C-terminally fused with mScarlet. The S84A mutation prevents S84 phosphorylation.
248273pHR TRAK2 S84E fl mScarlet Encodes TRAK2 S84E fl C-terminally fused with mScarlet. The S84E mutation mimics S84 phosphorylation.
248274pHR TRAK2 F92A, M95A, d373-380 fl mScarlet Encodes TRAK2 F92A, M95A, d373-380 fl C-terminally fused with mScarlet.
248275pHR dMiro (1-619) m-sfGFP FLAG I3-01Encodes the cytosolic portion of Drosophila Miro (dMiro) C-terminally fused with monomerized sfGFP (V206R), FLAG and the self-assembling I3-01 domain.
248276pHR Milton (1-755) mScarlet Encodes Milton (1-755) C-terminally fused with mScarlet. Expression of this fragment facilitates Milton-mediated motility of dMiro decorated nanocages in human cells.
248277pHR FRB m-sfGFP FLAG GB1 I3-01Encodes FRB C-terminally fused with monomerized sfGFP (V206R), FLAG and the self-assembling I3-01 domain.

Antibodies from Article