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OpenPlant Kit
(Kit # 1000000272 )

Depositing Lab:   Jim Haseloff

The OpenPlant toolkit is a collection of standardized DNA parts and vectors, for genetic engineering of the nuclear and plastid genomes of the liverwort Marchantia polymorpha, an emerging plant synthetic biology model. Although primarily developed for Marchantia, the toolkit can also be used in other plants such as the angiosperm Arabidopsis thaliana and the hornwort Anthoceros agrestis.

This kit will be sent as bacterial glycerol stocks in 96-well plate format.

Original Publication

Systematic Tools for Reprogramming Plant Gene Expression in a Simple Model, Marchantia polymorpha. Sauret-Güeto S, Frangedakis E, Silvestri L, Rebmann M, Tomaselli M, Markel K, Delmans M, West A, Patron NJ, Haseloff J. ACS Synth Biol. 2020 Apr 17;9(4):864-882. doi: 10.1021/acssynbio.6b00337. PubMed (Link opens in a new window) Article (Link opens in a new window)

Construction of DNA Tools for Hyperexpression in Marchantia Chloroplasts. Frangedakis E, Guzman-Chavez F, Rebmann M, Markel K, Yu Y, Perraki A, Tse SW, Liu Y, Rever J, Sauret-Gueto S, Goffinet B, Schneider H, Haseloff J. ACS Synth Biol. 2021 Jul 16;10(7):1651-1666. doi: 10.1021/acssynbio.0c00637. PubMed (Link opens in a new window) Article (Link opens in a new window)

Description

The OpenPlant toolkit is based on Loop assembly (Pollack et al., 2018 (Link opens in a new window)), a Golden Gate cloning method for DNA construct generation that employs Type IIS restriction enzymes and a recursive strategy to greatly simplify the process of plasmid assembly. It allows rapid and efficient production of large DNA constructs from DNA parts that follow a common assembly syntax. Unlike other systems that require elaborate sets of vectors, Loop assembly requires only two sets of four complementary vectors. In a series of reactions, standardized DNA parts can be assembled into multi-transcriptional units.

The OpenPlant toolkit was deposited by Jim Haseloff’s lab and is a collection of 79 plasmids including Loop nuclear transformation vectors, Loop vectors for chloroplast transformation, Loop vectors for CRISPR genome editing, and a series of standardized DNA L0 parts (promoters, 5' untranslated regions, signal peptides, coding sequences, and terminators) for expression in Marchantia. pCk and Loop vectors for CRISPR genome editing are supplied as individual stabs (please see Accessory Plasmids tab).

Panel A depicts the layout of DNA part positions and fusion sites (described in the figure caption). The DNA parts are labeled according to the Phytobrick common syntax (A1 to A3, B1 to B6, and C1). From left to right, the fusion sites and DNA parts are: GGAG, distal promoter (A1), TGAC, proximal promoter (A2–A3), TACT, 5’ UTR (B1–B2), AATG, CDS1 (B3), GCAGCC, CDS2 (B4), GCTTCG, CTAG (B5), GCTT, 3’ UTR (B6), GGTA, terminator (C1), CGCT. Panel B depicts pUAP4, containing a LacZ coding region flanked by SapI and BsaI cut sites. Above the plasmid is a DNA part, flanked by parts fusion sites and SapI sites. Using SapI and T4 ligase, the DNA part is placed in pUAP4, creating a level 0 plasmid. The BsaI sites can then be used to assemble transcriptional units (level 1 plasmids). Panel C depicts the pCk and pCs vectors for nuclear and chloroplast applications, along with the Ck and Cs plasmids for CRISPR/Cas9. All plasmids contain the black circles for pSa origin and blue rounded rectangles for lacZa cassette. Only the pCk, pCs, and CRISPR plasmids (not chloroplast plasmids) contain left and right border repeats.

Figure 1: Key elements in the OpenPlant Loop assembly toolkit. (A) Schematic of the parts and common syntax used in OpenPlant Loop assembly. A transcriptional unit is divided into a 5' non-transcribed region, a transcribed region, and a 3' non-transcribed region. These parts can be further broken down into the 5' UTR, the distal (PROMD) and proximal (PROMP) promoters, CDS12 (which can be split into CDS1 and CDS2), CTAG, 3' UTR, and the terminator (TERM). (B) Schematic of the pUAP4 vector containing SapI sites to accept L0 parts and BsaI sites to assemble L0 parts into transcription units (L1). (C) Summary of the Loop acceptor vectors of the OpenPlant toolkit. For nuclear genome transformation: pCk (1, 2, 3, and 4) assembles L0 parts into a Level 1 plasmid using BsaI, and pCs (A,B,C,E) assembles up to four Level 1 plasmids into a Level 2 construct using SapI. For chloroplast applications, pCkchlo (1, 2, 3, and 4) and pCschlo (A, B, C, and E) can be used for assembly. L1_lacZgRNA-Ck2, L1_lacZgRNA-Ck3, and L2_lacZgRNA-Cas9 are designed for CRISPR/Cas9 genome editing. LB and RB: left and right border repeats from nopaline C58 T-DNA. Filled blue rounded rectangle: lacZα cassette for blue-white screening. Filled black circles: pSa origin of replication. Image and caption derived from Figure 2 in Sauret-Güeto et al., 2020 (Link opens in a new window).

DNA parts are shown under the heading “Transcriptional Unit”, and are further split into three categories: promoter (PROM) plus 5’ UTR, CDS, and 3’ UTR plus terminator (TERM). PROM plus 5’ UTR can be split into individual parts or combined. Combined PROM and 5’ UTR parts include: MpEF1a5U, 35S, 35S2x, MpU6, PROM5_Mprpo_chlo, PROM5_NtpsbA_chlo, PROM5_Ntprrn_rev_chlo, and PROM5_MprbcL_short_chlo. PROM parts include: MpUBE2, MpNOP1, and NtpsbA_chlo. 5’ UTR parts include: MpUBE2, MpNOP1, Mppetb_short_chlo, and MprbcL_chlo. The CDS can be split into individual parts (CDS12 and CTAG) or combined. Combined parts include: hptII(HygR), mALS(CsR), nptII(KanR), GUS, LUC, AtcoCas9-NLS, CDS_aadA_chlo, and CDS_mTurquoise2_chlo. CDS12 parts include: eGFP, eGFP-linker, mVenus, mVenus-linker, mTurquoise2, mTurquoise2-linker, mScarlet-I, mScarlet-I-linker, MpER-Targ, TP-MpSIG2, and AtcoCas9-NLS. CTAG parts include eGFP, linker-eGFP, mVenus, linker-mVenus, mTurquoise2, linker-mTurquoise2, mScarlet-I, eGFP-HDEL, eGFP-HnGnHDEL, mVenus-HDEL, mTurquoise2-HDEL, linker-N7, NLS-SV40, and linker-Lti6b. 3’ UTR and TERM parts include: 35S, nos, nos-35S, MpNOP1, 3TERM_NtpsbA_rev_chlo, and 3TERM_BBa_B0015_chlo.

Figure 2: OpenPlant Kit DNA parts. Image modified from Sauret-Güeto et al., 2020 (Link opens in a new window).

Kit Documentation

OpenPlant protocols are available at:

How to Cite this Kit

These plasmids were created by your colleagues. Please acknowledge the Principal Investigator, cite the article in which they were created, and include Addgene in the Materials and Methods of your future publications.

For your Materials and Methods section:

"The OpenPlant Kit was a gift from Jim Haseloff (Addgene kit #1000000272)."

For your Reference section:

Systematic Tools for Reprogramming Plant Gene Expression in a Simple Model, Marchantia polymorpha. Sauret-Güeto S, Frangedakis E, Silvestri L, Rebmann M, Tomaselli M, Markel K, Delmans M, West A, Patron NJ, Haseloff J. ACS Synth Biol. 2020 Apr 17;9(4):864-882. doi: 10.1021/acssynbio.6b00337. PubMed (Link opens in a new window) Article (Link opens in a new window)

Construction of DNA Tools for Hyperexpression in Marchantia Chloroplasts. Frangedakis E, Guzman-Chavez F, Rebmann M, Markel K, Yu Y, Perraki A, Tse SW, Liu Y, Rever J, Sauret-Gueto S, Goffinet B, Schneider H, Haseloff J. ACS Synth Biol. 2021 Jul 16;10(7):1651-1666. doi: 10.1021/acssynbio.0c00637. PubMed (Link opens in a new window) Article (Link opens in a new window)

OpenPlant kit - #1000000272

Resistance Color Key

Each circle corresponds to a specific antibiotic resistance in the kit plate map wells.

Inventory

Searchable and sortable table of all plasmids in kit. The Well column lists the plasmid well location in its plate. The Plasmid column links to a plasmid's individual web page.

Kit Plate Map

96-well plate map for plasmid layout. Hovering over a well reveals the plasmid name, while clicking on a well opens the plasmid page.

Resistance Color Key

Spectinomycin
Kanamycin
Chloramphenicol

Inventory

Well Plasmid Resistance
A / 1 pCsA
Spectinomycin
A / 2 pCsB
Spectinomycin
A / 3 pCsC
Spectinomycin
A / 4 pCsE
Spectinomycin
A / 5 pCk1_spacer
Kanamycin
A / 6 pCk2_spacer
Kanamycin
A / 7 pCk3_spacer
Kanamycin
A / 8 pCk4_spacer
Kanamycin
A / 9 pCsA_spacer
Spectinomycin
A / 10 pCsB_spacer
Spectinomycin
A / 11 pCsC_spacer
Spectinomycin
A / 12 pCsE_spacer
Spectinomycin
B / 1 pUAP4
Chloramphenicol
B / 2 CDS_nptII
Chloramphenicol
B / 3 CDS_mALS
Chloramphenicol
B / 4 CDS_hptII
Chloramphenicol
B / 5 CDS_GUS
Chloramphenicol
B / 6 CDS12_mScarlet-I
Chloramphenicol
B / 7 CDS12_mScarletI-linker
Chloramphenicol
B / 8 CDS12_mTurquoise2
Chloramphenicol
B / 9 CDS12_mTurquoise2-linker
Chloramphenicol
B / 10 CDS12_mVenus
Chloramphenicol
B / 11 CDS12_mVenus-linker
Chloramphenicol
B / 12 CDS12_TP-MpSIG2
Chloramphenicol
C / 1 CDS12-MpER-Targ
Chloramphenicol
C / 2 CTAG_linker-Lti6b
Chloramphenicol
C / 3 CTAG_linker-mTurquoise2
Chloramphenicol
C / 4 CTAG_linker-mVenus
Chloramphenicol
C / 5 CTAG_linker-N7
Chloramphenicol
C / 6 CTAG_mScarlet-I
Chloramphenicol
C / 7 CTAG_mTurquoise2
Chloramphenicol
C / 8 CTAG_mTurquoise2-HDEL
Chloramphenicol
C / 9 CTAG_mVenus
Chloramphenicol
C / 10 CTAG_mVenus-HDEL
Chloramphenicol
C / 11 CTAG_NLS-SV40
Chloramphenicol
C / 12 PROM_MpUBE2
Chloramphenicol
D / 1 5UTR_MpUBE2
Chloramphenicol
D / 2 PROM5_35S
Chloramphenicol
D / 3 PROM5_35Sx2
Chloramphenicol
D / 4 PROM5_MpEF1a
Chloramphenicol
D / 5 3TERM_35S
Chloramphenicol
D / 6 3TERM_Nos
Chloramphenicol
D / 7 3TERM_Nos-35S
Chloramphenicol
D / 8 PROM_MpNOP1
Chloramphenicol
D / 9 5UTR_MpNOP1
Chloramphenicol
D / 10 3TERM_MpNOP1
Chloramphenicol
D / 11 L1_KanR-Ck1
Kanamycin
D / 12 L1_CsR-Ck1
Kanamycin
E / 1 L1_HyR-Ck1
Kanamycin
E / 2 L1_NOP1:mS-N7-Ck2
Kanamycin
E / 3 L1_UBE2:mT-N7-Ck2
Kanamycin
E / 4 L1_UBE2:mT-N7-Ck3
Kanamycin
E / 5 L1_UBE2:mS-Lt-Ck3
Kanamycin
E / 6 L1_UBE2:mS-Lt-Ck4
Kanamycin
E / 7 L2_UBE2:mT-N7-CsA
Spectinomycin
E / 8 pCk1chlo
Kanamycin
E / 9 pCk2chlo
Kanamycin
E / 10 pCk3chlo
Kanamycin
E / 11 pCk4chlo
Kanamycin
E / 12 pCsAchlo
Spectinomycin
F / 1 pCsBchlo
Spectinomycin
F / 2 pCsCchlo
Spectinomycin
F / 3 pCsEchlo
Spectinomycin
F / 4 PROM_Ntpsba_chlo
Chloramphenicol
F / 5 PROM5_Mprnpo_chlo
Chloramphenicol
F / 6 PROM5_Ntprrn_rev_chlo
Chloramphenicol
F / 7 PROM5_Ntpsba_chlo
Chloramphenicol
F / 8 PROM5_MprbcL_short_chlo
Chloramphenicol
F / 9 3TERM_bba_b0015_rev_chlo
Chloramphenicol
F / 10 3TERM_Ntpsba_rev_chlo
Chloramphenicol
F / 11 5UTR_Mppetb_short_chlo
Chloramphenicol
F / 12 5UTR_Mprbcl_chlo
Chloramphenicol
G / 1 CDS_aadA_chlo
Chloramphenicol
G / 2 CDS_mTurqcp_chlo
Chloramphenicol
G / 3 L1-Ck1-hs1-l-chlo
Kanamycin
G / 4 L1-Ck1-hs2-l-chlo
Kanamycin
G / 5 L1-Ck2-smR-chlo
Kanamycin
G / 6 L1-Ck4-hs1-r-chlo
Kanamycin
G / 7 L1-Ck4-hs2-r-chlo
Kanamycin
Data calculated @ 2026-02-07

Kit Plate Map - #1000000272

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