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Fluorescent Proteins: Subcellular Localization


Background

The subcellular localization of a protein is often tied to its function, so it's important to determine where your protein of interest resides. There are a variety of methods for determining subcellular localization. One of the most common is to use fluorescence microscopy to compare the localization of your protein of interest with known markers.

Addgene has assembled a collection of fluorescent fusion proteins that are targeted to various organelles or subcellular structures. You should fuse your protein of interest with a fluorescent protein that has a different emission wavelength. For example, if the marker protein is fused to GFP, you may want to fuse your protein to a red fluorescent protein (see this fusion protein guide to find the appropriate backbone). Then, by examining the overlap between your fusion protein and the marker fusion protein using fluorescence microscopy, you can assess whether your protein is targeted to the same structures as these well-characterized proteins.


General Structures

The cell contains a variety of organelles and subcellular structures each with their own unique function. The nucleus, for example, houses the cell’s genetic material. The mitochondria produces the energy cells required for chemical reactions. Labeling these structures can be quite useful for identifying the localization or possible function of a protein of interest. Browse or search the table below to find markers of general cellular organelles and structures.

ID Plasmid Structure Targeting Gene/Signal Fluorescent Protein PI
98875 3xnls-mNeonGreen Nucleus 3xNLS mNeonGreen Dorus Gadella
98817 3xnls-mTurquoise2 Nucleus 3xNLS mTurquoise2 Dorus Gadella
98816 3xnls-mScarlet-I Nucleus 3xNLS mScarlet-I Dorus Gadella
98876 4xmts-mNeonGreen Mitochondria 4xmts mNeonGreen Dorus Gadella
98819 4xmt-mTurquoise2 Mitochondria 4xmts mTurquoise2 Dorus Gadella
98818 4xmts-mScarlet-I Mitochondria 4xmts mScarlet-I Dorus Gadella
19031 pABCb10-GFP Mitochondria ABCb10 GFP Orian Shirihai
158003 pCMV-mGold-CAF1-C-10 Nucleus CAF-1 mGold Francois St-Pierre
27705 CAV1-mCherry Caveolae Cav1 mCherry Ari Helenius
37533 cfSGFP2-N Extracellular milieu cfSGFP2-N EGFP Ikuo Wada
73208 pcDNA3.1-COXIV-COX8-dL5-2XG4S-mCer3 Mitochondria COX IV-derived (1-22 aa) import sequence and COX VIII signal peptide dL5 FAP; mCerulean3 Marcel Bruchez
44385 pLV-mitoGFP Mitochondria COX8 GFP Pantelis Tsoulfas
80000 pMito-miRFP703 Mitochondria COX8A miRFP703 Vladislav Verkhusha
36208 pmTurquoise2-Mito Mitochondria COX8A(1-29) mTurquoise2 Dorus Gadella
26673 pEGFP-C1-Fibrillarin Nucleolus Fibrillarin EGFP Sui Huang
36207 pmTurquoise2-H2A Nucleus H2A mTurquoise2 Dorus Gadella
85053 pmScarlet-i_H2A_C1 Nucleus H2A mScarlet-I Dorus Gadella
85052 pmScarlet-H_H2A_C1 Nucleus H2A mScarlet-H Dorus Gadella
85051 pmScarlet_H2A_C1 Nucleus H2A mScarlet Dorus Gadella
18982 pHIV-H2BmRFP Chromatin H2B mRFP Bryan Welm, Zena Werb
21045 pH2B_mCherry_IRES_puro2 Chromatin H2B mCherry Daniel Gerlich
25999 LV-GFP Chromatin H2B GFP Elaine Fuchs
11680 H2B-GFP Chromatin H2B GFP Geoff Wahl
21210 PGK-H2BeGFP Chromatin H2B EGFP Mark Mercola
21217 PGK-H2BmCherry Chromatin H2B mCherry Mark Mercola
20972 H2B-mCherry Chromatin H2B mCherry Robert Benezra
80001 pH2B-miRFP703 Chromatin H2B miRFP703 Vladislav Verkhusha
17650 GFP-HP1gamma Chromatin HP1 gamma GFP Tom Misteli
55068 mCherry-LaminA-C-18 Nuclear Envelope LaminA1 mCherry* Michael Davidson
98831 mScarlet-I-LaminB Nuclear Envelope LaminB mScarlet-I Dorus Gadella
98830 mTurquoise2-LaminB Nuclear Envelope LaminB mTurquoise2 Dorus Gadella
55069 mCherry-LaminB1-10 Nuclear Envelope LaminB1 mCherry* Michael Davidson
98822 Lck-mTurquoise2 Plasma Membrane Lck N-terminal lipidation signal mTurquoise2 Dorus Gadella
98821 Lck-mScarlet-I Plasma Membrane Lck N-terminal lipidation signal mScarlet-I Dorus Gadella
50057 pLYS1-FLAG-MitoGFP-HA Mitochondria MCU GFP Vamsi Mootha
49153 GFP-Mff Mitochondria-Outer Membrane Mff acGFP1 Gia Voeltz
158007 pCMV-mGold-Mito-N-7 Mitochondria Cox8A mGold Francois St-Pierre
49151 mito-BFP Mitochondria mitochondrial targeting signal (COX4) TagBFP Gia Voeltz
55102 mCherry-Mito-7 Mitochondria Mitochondrial targeting sequence(COX8A) mCherry* Michael Davidson
37354 pQC NLS mCherry IX Nucleus NLS mCherry Connie Cepko
37341 pQC NLS YFP IX Nucleus NLS YFP Connie Cepko
158000 pCaggs-NLS-PAmCherry1-GSS-EGFP Nucleus NLS mGold Francois St-Pierre
14893 pEBFP2-Nuc Nucleus NLS EBFP2 Robert Campbell
14894 pmKalama1-Nuc Nucleus NLS mKamala1 Robert Campbell
73205 pcDNA3.1-NLS-myc-dL5-2xG4S-mCer3 Nucleus NLS (from Mak16p protein) dL5 FAP; mCerulean3 Marcel Bruchez
73207 pcDNA3.1-KozATG-dL5-2XG4S-mCer3 Cytosol, nucleoplasm None dL5 FAP; mCerulean3 Marcel Bruchez
36206 pmTurquoise2-NES Non-nucleus Nuclear Export Sequence mTurquoise2 Dorus Gadella
85062 pmScarlet-i_NES_C1 Non-nucleus Nuclear Export Sequence mScarlet-I Dorus Gadella
85061 pmScarlet-H_NES_C1 Non-nucleus Nuclear Export Sequence mScarlet-H Dorus Gadella
85060 pmScarlet_NES_C1 Non-nucleus Nuclear Export Sequence mScarlet Dorus Gadella
37351 pQC membrane TdTomato IX Membrane Palmitoylation TdTomato Connie Cepko
22479 FUmGW Membrane Palmitoylation GFP Connie Cepko
14757 pCAG-mGFP Membrane Palmitoylation sequence from GAP43 EGFP Connie Cepko
61099 Lck-GFP Membrane Palmitoylation sequence from Lck EGFP Steven Green
36209 pPalmitoyl-mTurquoise2 Membrane Palmitoylation sequence from p63 mTurquoise2 Dorus Gadella
73206 pcDNA3.1-kappa-myc-dL5-2xG4S-TMst Cell surface (mammalian) PDGFR-derived dL5 FAP; mCerulean3 Marcel Bruchez
45944 pTDpelB-C_sfYFPTwinStrep Periplasmic space PelB signal sequence YFP Thorben Dammeyer
54520 mCherry-Peroxisomes-2 Peroxisome Peroximal Targeting Signal 1 (PTS1) mCherry* Michael Davidson
36203 pmTurquoise2-Peroxi Peroxisome Peroxisomal Targeting Sequence mTurquoise2 Dorus Gadella
17990 pEGFP-SF2 Nucleus SF2 EGFP Tom Misteli
137806 mNeonGreen-peroxi Peroxisome SKL mNeonGreen Dorus Gadella
73210 pcDNA3.1-KozATG-myc-dL5-2XG4S-mCer3-SKL Peroxisome SKL tripeptide, peroxisome transport signal dL5 FAP; mCerulean3 Marcel Bruchez
85065 pmScarlet-I_peroxisome_C1 Peroxisome SRL mScarlet-I Dorus Gadella
85064 pmScarlet-H_peroxisome_C1 Peroxisome SRL mScarlet-H Dorus Gadella
85063 pmScarlet_peroxisome_C1 Peroxisome SRL mScarlet Dorus Gadella
55146 mCherry-TOMM20-N-10 Mitochondria-Outer Membrane TOMM20 mCherry* Michael Davidson

*Fusions to other FPs can be found by searching the complete Michael Davidson Fluorescent Protein Collection.

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Cytoskeletal Elements

The cytoskeleton is the cell’s internal framework that assists in sustaining structure and function. The cytoskeleton is primarily made up of microtubules, actin filaments, and intermediate filaments. These structures give the cell its shape, organize cellular organelles, and are necessary for movement and cell division. Browse or search the table below to find markers of cytoskeletal element.

ID Plasmid Structure Targeting Gene/Signal Fluorescent Protein PI
26722 GFP-cortactin Actin Filaments Cortactin EGFP Anna Huttenlocher
32907 pEGFP-rNFM Intermediate Filaments (neuronal) Nefm EGFP Anthony Brown
32909 pEGFP-mNFM Intermediate Filaments (neuronal) Nefm EGFP Anthony Brown
11908 pEGFP-N1 alpha-actinin 1 Actin Filaments alpha-actinin 1 EGFP Carol Otey
27676 Cortactin-pmCherryC1 Actin Filaments Cortactin mCherry Christien Merrifield
36201 pLifeAct-mTurquoise2 Actin Filaments LifeAct mTurquoise2 Dorus Gadella
36202 pmTurquoise2-Tubulin Microtubules alpha-tubulin mTurquoise2 Dorus Gadella
85045 pmScarlet_alphaTubulin_C1 Microtubules alpha-tubulin mScarlet Dorus Gadella
85046 pmScarlet-H_alphaTubulin_C1 Microtubules alpha-tubulin mScarlet-H Dorus Gadella
85047 pmScarlet-i_alphaTubulin_C1 Microtubules alpha-tubulin mScarlet-I Dorus Gadella
85054 pLifeAct_mScarlet_N1 Actin Filaments LifeAct mScarlet Dorus Gadella
85055 pLifeAct_mScarlet-H_N1 Actin Filaments LifeAct mScarlet-H Dorus Gadella
85056 pLifeAct_mScarlet-i_N1 Actin Filaments LifeAct mScarlet-I Dorus Gadella
137801 EMTB-mScarletI Microtubules EMTB mScarletI Dorus Gadella
137802 EMTB-mNeonGreen Microtubules EMTB mNeonGreen Dorus Gadella
137803 EMTB-mTurquoise2 Microtubules EMTB mTurquoise2 Dorus Gadella
137807 mTurquoise2-MapTau Microtubules MapTau mTurquoise2 Dorus Gadella
137808 mNeonGreen-MapTau Microtubules MapTau mNeonGreen Dorus Gadella
137811 ITPKA-mTurquoise2 Actin Filaments ITPKA mTurquoise2 Dorus Gadella
58473 pEGFP-C1 F-tractin-EGFP Actin Filaments F-tractin EGFP Dyche Mullins
41147 pEGFP Centrin2 Centrioles (dependent on cell cycle) Centrin-2 EGFP Erich Nigg
41151 pEGFP Cep170 C-term Centrioles (dependent on cell cycle) Cep170 EGFP Erich Nigg
41163 pEGFP PICH Centromeres (dependent on cell cycle) PICH EGFP Erich Nigg
158008 pCMV-mGold-Tubulin-C-18 Microtubules alpha-tubulin mGold Francois St-Pierre
158009 pCMV-mGold-Actin-C-18 Actin Filaments Actin mGold Francois St-Pierre
49149 mCh-alpha-tubulin Microtubules alpha-tubulin mCherry Gia Voeltz
89472 GFP-hChibby1 Mother Centrioles / Ciliary Base Chibby1 GFP Ken-Ichi Takemaru
89770 pLenti-EGFP-hChibby1 Mother Centrioles / Ciliary Base Chibby1 EGFP Ken-Ichi Takemaru
118084 pLenti-EB1-EGFP Microtubules EB1 EGFP Ken-Ichi Takemaru
122871 GFP-hCCDC11 Centriolar satellites CCDC11 EGFP Ken-Ichi Takemaru
54234 mEmerald-PLK1-N-16 Centrosome Plk1 mEmerald* Michael Davidson
54491 mCherry-Lifeact-7 Actin Filaments LifeAct mCherry* Michael Davidson
54610 mEGFP-Lifeact-7 Actin Filaments LifeAct EGFP Michael Davidson
55148 mCherry-Tubulin-C-18 Microtubules alpha-tubulin mCherry* Michael Davidson
12298 pIRESneo-EGFP-alpha Tubulin Microtubules alpha-tubulin EGFP Patricia Wadsworth
27123 eTC GFP beta-actin full length Actin Filaments beta-actin EGFP Robert Singer
27382 pDEST/N1-hEB1-GFP Microtubules EB1 GFP Robin Shaw
40908 pDEST/LIfeAct-mCherry-N1 Actin Filaments LifeAct mCherry Robin Shaw
21948 pCAG-mGFP-Actin Actin Filaments beta-actin EGFP Ryohei Yasuda
31930 pPAmCherry-a-tubulin Microtubules alpha-tubulin PAmCherry Vladislav Verkhusha
31949 pPAmCherry-b-actin Actin Filaments beta-actin PAmCherry Vladislav Verkhusha
79990 pKeratin-miRFP703 Intermediate Filaments (epithelial) Keratin miRFP703 Vladislav Verkhusha
79991 pmiRFP703-Tubulin Microtubules alpha-tubulin miRFP703 Vladislav Verkhusha
79993 pLifeAct-miRFP703 Actin Filaments LifeAct miRFP703 Vladislav Verkhusha
79994 pEB3-miRFP703 Microtubules EB3 miRFP703 Vladislav Verkhusha
79996 pVimentin-miRFP703 Intermediate Filaments (vimentin) Vimentin miRFP703 Vladislav Verkhusha
26737 GFP-UtrCH Actin Filaments Utrophin (aa# 1-261) GFP William Bement
26740 mCherry-UtrCH Actin Filaments Utrophin mCherry William Bement
26741 EMTB-3XGFP Microtubules ensconsin GFP William Bement
26742 EMTB-mCherry Microtubules ensconsin mCherry William Bement

*Fusions to other FPs can be found by searching the complete Michael Davidson Fluorescent Protein Collection.

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Cell Junctions

Cell junctions are responsible for forming connections and adhesions between animal cells. They are important for holding cell structures together and reducing cell stress. There are four types of cell junctions: adherens junctions, gap junctions, tight junctions, and desmosomes. Browse or search the table below to find markers for cellular junctions.

ID Plasmid Structure Targeting Gene/Signal Fluorescent Protein PI
71367 Human Beta-catenin GFP Adherens Junctions Beta-catenin EGFP Alpha Yap
67937 mouse E-cadherin GFP (423) Adherens Junctions E-cadherin EGFP Alpha Yap
71366 Murine E-cadherin mCherry Adherens Junctions E-cadherin mCherry Alpha Yap
54219 mEmerald-Paxillin-22 Focal Adhesions paxillin mEmerald* Michael Davidson
15233 Paxillin-pEGFP Focal Adhesions paxillin GFP Rick Horwitz
26720 RFP-zyxin Focal Adhesions Zyxin RFP Anna Huttenlocher
11908 pEGFP-N1 alpha-actinin 1 Focal Adhesions alpha-actinin 1 EGFP Carol Otey
31923 pPaxillin-PSmOrange Focal Adhesions paxillin PSmOrange Vladislav Verkhusha
79999 pZyxin-miRFP703 Focal Adhesions Zyxin miRFP703 Vladislav Verkhusha
55023 mCherry-Cx43-7 Gap Junctions Connexin 43 / GJA1-20k mCherry* Michael Davidson
40907 pDEST/hCx43-EGFP-N1 Gap Junctions Connexin 43 EGFP Robin Shaw
49861 pDEST-mCherry-GJA1-20k-N1 Gap Junctions Connexin 43 / GJA1-20k mCherry Robin Shaw
69007 Cx43-msfGFP Gap Junctions Connexin 43 / GJA1 sfGFP David Spray
54959 mApple-VE-Cadherin-N-10 Tight Junctions VE-Cadherin mApple Michael Davidson
55165 mCherry-ZO1-C-14 Tight Junctions Zonula Occludens-1 mCherry Michael Davidson
55001 mCherry-Beta-Catenin-20 Adherens Junctions Beta-Catenin mCherry Michael Davidson
30313 pEGFP ZO1 Tight Junctions Zonula Occludens-1 GFP Alan Fanning

*Fusions to other FPs can be found by searching the complete Michael Davidson Fluorescent Protein Collection.

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Secretory Pathway

The secretory pathway is named for its role in secreting proteins from the cell to the extracellular environment. Proteins are first translated at the endoplasmic reticulum, then are transported to the Golgi apparatus in vesicles to be packaged and shipped. Many of the proteins that enter the secretory pathway require chaperones to help with folding or post-translational modification to function properly. Browse or search the table below to find markers of secretory pathway components.

ID Plasmid Structure Targeting Gene/Signal Fluorescent Protein PI
110060 FUW mCherry-GFP-LC3 Autophagosome LC3 mCherry, GFP Anne Brunet
40827 mCherry-hLC3B-pcDNA3.1 Autophagosome LC3 mCherry David Rubinsztein
11546 EGFP-LC3 Autophagosome LC3 EGFP Karla Kirkegaard
61205 pCMB-AUTr Autophagosome LC3(ATG8) mCherry Maria Harrison
38277 pMXs-puro GFP-p62 Autophagosome Sequestosome-1 EGFP Noboru Mizushima
38272 pMXs-IP GFP-WIPI-1 Autophagosome WIPI1 EGFP Noboru Mizushima
21074 ptfLC3 Autophagosome LC3 EGFP Tamotsu Yoshimori
21073 pEGFP-LC3 Autophagosome LC3 EGFP Tamotsu Yoshimori
24920 pEGFP-LC3 (human) Autophagosome LC3 EGFP Toren Finkel
14435 mRFP-Clc Clathrin-coated vesicles Clathrin, light polypeptide mRFP Ari Helenius
70217 pEGFP-GFP11-Clathrin light chain Clathrin-coated vesicles Clathrin light chain GFP Bo Huang
54040 mEmerald-Clathrin-15 Clathrin-coated vesicles Clathrin mEmerald* Michael Davidson
79997 pmiRFP703-Clathrin Clathrin-coated vesicles Clathrin miRFP703 Vladislav Verkhusha
21741 Clathrin-LCa-EYFP Clathrin-coated vesicles Clathrin, light polypeptide EYFP Xiaowei Zhuang
14437 mRFP-Rab5 Early endosomes RAB5A mRFP Ari Helenius
61801 GFP-Rab4B Early endosomes Rab4B AcGFP Gia Voeltz
49201 mCh-Rab5 Early endosomes Rab5 mCherry Gia Voeltz
49147 BFP-Rab5 Early endosomes Rab5 BFP Gia Voeltz
61802 GFP-Rab5B Early endosomes Rab5B AcGFP Gia Voeltz
79802 pTag-RFP-C-h-Rab5a-c-Myc Early endosomes Rab5a TagRFP James Johnson
79801 pTag-BFP-C-h-Rab5a-c-Myc Early endosomes Rab5a TagBFP James Johnson
13050 DsRed-Rab5 WT Early endosomes RAB5A DsRed2 Richard Pagano
42307 GFP-EEA1 wt Early endosomes EEA1 GFP Silvia Corvera
42635 TagRFP-T-EEA1 Early endosomes EEA1 TagRFP-T Silvia Corvera
38770 pEF.myc.ER-E2-Crimson Endoplasmic Reticulum ER retention signal E2-Crimson Benjamin Glick
36204 pmTurquoise2-ER Endoplasmic Reticulum ER retention signal mTurquoise2 Dorus Gadella
137804 ER-mNeonGreen Endoplasmic Reticulum KDEL sequence mNeonGreen Dorus Gadella
137805 ER-mScarletI Endoplasmic Reticulum KDEL sequence mScarlet-I Dorus Gadella
68072 ERmoxGFP Endoplasmic Reticulum ER retention signal moxGFP Erik Snapp
68126 ERoxBFP Endoplasmic Reticulum ER retention signal oxBFP Erik Snapp
49150 BFP-KDEL Endoplasmic Reticulum BiP ER signal sequence BFP Gia Voeltz
61807 Rtn4a-GFP Endoplasmic Reticulum Rtn4a AcGFP Gia Voeltz
49155 mCh-Sec61 beta Endoplasmic Reticulum Sec61 mCherry Gia Voeltz
49154 BFP-Sec61 beta Endoplasmic Reticulum Sec61 BFP Gia Voeltz
73209 pcDNA3.1-kappa-myc-dL5-2XG4S-mCer3-KDEL Endoplasmic Reticulum KDEL dL5 FAP; mCerulean3 Marcel Bruchez
73209 pcDNA3.1-kappa-myc-dL5-2XG4S-mCer3-KDEL Endoplasmic Reticulum KDEL dL5 FAP; mCerulean3 Marcel Bruchez
85068 pCytERM_mScarlet-i_N1 Endoplasmic Reticulum CytERM mScarlet-I Dorus Gadella
85067 pCytERM_mScarlet-H_N1 Endoplasmic Reticulum CytERM mScarlet-H Dorus Gadella
85066 pCytERM_mScarlet_N1 Endoplasmic Reticulum CytERM mScarlet Dorus Gadella
158004 pCMV-mGold-Calnexin-N-14 Endoplasmic Reticulum Calnexin mGold Francois St-Pierre
158002 pCMV-mGold-Rab4a-C-7 Endosomes Rab4a mGold Francois St-Pierre
14873 GolgiCFP Golgi apparatus eNOS(1-33) CFP Alexandra Newton
36205 pmTurquoise2-Golgi Golgi apparatus B4GALT1(1-61) mTurquoise2 Dorus Gadella
68073 GalToxBFP Golgi apparatus GalT signal anchor transmembrane domain oxBFP Erik Snapp
158005 pCMV-mGold-MannII-N-10 Golgi apparatus MannII mGold Francois St-Pierre
85050 pmScarlet-i_Giantin_C1 Golgi apparatus Giantin mScarlet-I Dorus Gadella
85049 pmScarlet-H_Giantin_C1 Golgi apparatus Giantin mScarlet-H Dorus Gadella
85048 pmScarlet_Giantin_C1 Golgi apparatus Giantin mScarlet Dorus Gadella
129630 mTurquoise2-Giantin Golgi apparatus Giantin mTurquoise2 Dorus Gadella
98880 mNeonGreen-Giantin Golgi apparatus Giantin mNeonGreen Dorus Gadella
61804 mCh-Rab7A Late endosomes Rab7a mCherry Gia Voeltz
61803 GFP-Rab7A Late endosomes Rab7a AcGFP Gia Voeltz
79804 pTag-RFP-C-h-Rab7-c-Myc Late endosomes Rab7 TagRFP James Johnson
79803 pTag-BFP-C-h-Rab7-c-Myc Late endosomes Rab7 TagBFP James Johnson
12605 GFP-rab7 WT Late endosomes RAB7 GFP Richard Pagano
12661 DsRed-rab7 WT Late endosomes RAB7 DsRed Richard Pagano
158006 pCMV-mGold-Lysosome-N-20 Lysosome Lamp1 mGold Francois St-Pierre
98882 Lamp1-mNeonGreen Lysosomes Lamp1 mNeonGreen Dorus Gadella
98828 Lamp1-mTurquoise2 Lysosomes Lamp1 mTurquoise2 Dorus Gadella
98827 Lamp1-mScarlet-I Lysosomes Lamp1 mScarlet-I Dorus Gadella
55073 mCherry-Lysosomes-20 Lysosomes Lamp1 mCherry* Michael Davidson
79998 pLAMP1-miRFP703 Lysosomes Lamp1 miRFP703 Vladislav Verkhusha
1816 Lamp1-YFP Lysosomes Lamp1 YFP Walther Mothes
1817 Lamp1-RFP Lysosomes Lamp1 RFP Walther Mothes
79806 pTag-RFP-C-h-Rab11a-c-Myc Recycling endosomes Rab11a TagRFP James Johnson
79805 pTag-BFP-C-h-Rab11a-c-Myc Recycling endosomes Rab11a TagBFP James Johnson
79800 pTag-RFP-C-h-Rab4a-c-Myc Recycling endosomes Rab4a TagRFP James Johnson
79799 pTag-BFP-C-h-Rab4a-c-Myc Recycling endosomes Rab4a TagBFP James Johnson
12674 GFP-rab11 WT Recycling endosomes Rab11a GFP Richard Pagano
12679 DsRed-rab11 WT Recycling endosomes Rab11a DsRed Richard Pagano

*Fusions to other FPs can be found by searching the complete Michael Davidson Fluorescent Protein Collection.

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